資源描述:
《探討水稻光周期開(kāi)花調(diào)控途徑》由會(huì)員上傳分享,免費(fèi)在線閱讀,更多相關(guān)內(nèi)容在行業(yè)資料-天天文庫(kù)。
1、http://www.tasysmm.comhttp://www.tahuayu.comhttp://www.talwhw.cnhttp://www.tskeda.cn探討水稻光周期開(kāi)花調(diào)控途徑開(kāi)花是許多高等植物生命周期循環(huán)的一個(gè)重要過(guò)程。植物開(kāi)花包括開(kāi)花決定、成花啟動(dòng)、花器官的發(fā)育[1]。植物能否成功地進(jìn)行有性繁殖取決于精確的開(kāi)花時(shí)間。開(kāi)花決定受到環(huán)境信號(hào)和內(nèi)源遺傳信號(hào)的控制,是開(kāi)花基因時(shí)空順序表達(dá)的結(jié)果。在這些信號(hào)中,最重要的一個(gè)決定因素是植物對(duì)于光周期的反應(yīng)。光周期指一日之內(nèi)晝夜長(zhǎng)度的相對(duì)變化,也即日長(zhǎng)。植物對(duì)于這種相對(duì)變化做出
2、的反應(yīng)稱為光周期現(xiàn)象。早在1852年,英國(guó)植物學(xué)家ArthurHenfrey就認(rèn)識(shí)到日長(zhǎng)對(duì)于開(kāi)花的重要性。20世紀(jì)20年代,Garner和Allard發(fā)現(xiàn)光照長(zhǎng)度對(duì)植物的生長(zhǎng)和發(fā)育,尤其是對(duì)決定植物開(kāi)花時(shí)間起著重要作用。根據(jù)對(duì)光周期的反應(yīng),一般可將植物分為3種:對(duì)長(zhǎng)日照敏感而成花的為長(zhǎng)日照植物;對(duì)短日照敏感而成花的為短日照植物;對(duì)日照長(zhǎng)度不敏感的為光期鈍感植物。擬南芥和水稻分別作為開(kāi)花時(shí)間調(diào)控研究的長(zhǎng)日照植物和短日照植物的模型。目前,對(duì)控制擬南芥開(kāi)花的光周期途徑,已經(jīng)有較為完整和深入的研究?,F(xiàn)結(jié)合擬南芥有關(guān)光周期開(kāi)花調(diào)控的研究進(jìn)展,就
3、有關(guān)決定模式生物水稻開(kāi)花時(shí)間的光周期調(diào)控的主要研究進(jìn)展作簡(jiǎn)要綜述。Blossomisanimportantprocessinmanyhigherplantlifecycle.Plantsblossomincludinghttp://www.tasysmm.comhttp://www.tahuayu.comhttp://www.talwhw.cnhttp://www.tskeda.cndevelopment[1]blossomdecided,floralevocation,floralorgan.Plantscansuccessfull
4、yreproducesexuallydependsontheblossomtimeaccurate.Blossomdecidediscontrolledbyenvironmentalsignalsandendogenousgeneticsignal,istheresultofexpressionofgenesequenceblossomtime.Inthesesignals,athemostimportantdeterminantisplantforphotoperiodresponse.Adaynightlightcycleleng
5、thvariation,arenowlong.Plantfortherelativechangemakethiskindofreactioniscalledphotoperiodism.Asearlyasin1852,theBritishbotanistArthurHenfreyrecognizedtheimportanceofthelengthofdaytoblossom.Nineteentwenties,GarnerandAllardfoundthatthelightlengthonplantgrowthanddevelopmen
6、t,especiallyforthedecisiontoblossomtimeplantplaysanimportantrole.Dependingonthephotoperiodresponse,thegeneralcanbedividedinto3kindsofplants:floweringoflong-daysensitivetolong-dayplant;toshortphotoperiodsensitiveandfloweringofshort-dayplant;onthedaylengthisnotsensitiveto
7、photoperiodhttp://www.tasysmm.comhttp://www.tahuayu.comhttp://www.talwhw.cnhttp://www.tskeda.cninsensitiveplants.Arabidopsisandricewereusedaslong-dayplantsblossomtimecontrolofshortdayplantmodel.Atpresent,tocontrolthephotoperiodpathwayofArabidopsisthalianablossom,thereha
8、vebeenmorecompleteandthoroughresearch.NowwiththeresearchprogressofArabidopsisphotoperiodregulationrelatedtoblo